In addition to the excellent answer by Poor, I can add from my own experience in astrophotography. When you want to attach your camera to a telescope to capture photos of deep sky objects, you'll need to know the exact focal point of the telescope objective. In a newtonian reflector, for example, the focal point/plane of the primary mirror falls somewhere in the focusing tube, and to get the image on the camera sensor, you'll attach the camera to the focuser using T-adapter and T-rings, and adjust it so that the sensor plane lies at the focal point of the primary mirror. Knowing the exact position of the sensor plane makes life much easier when you're trying to focus your telescope.